The elevation of the radiator or any other part of the cooling system is totally irrelevant. The tubes are there because there is no other way to get air out of the heads without them. Yes, you could bleed the air out of the tubes and then cap them, but you would have to do this every time you filled the cooling system. The air can not flow down into the block to get out. The tubes are at the top of the heads because that is where the air accumulates, and if you do not bleed it out, it will make the head run hot where the air is. When there is a pocket of air in the top of the head, then no water is in that section of the water jacket, why is that so hard to figure out? The engineers made an air trap in the heads of LS engines, but they resolved it with the bleed tubes, I would leave them in service. They did not put them in just for grins, they are needed to get the air out. Other manufacturers have had similar problems in other engines, and they have put in manual bleed valves to remove the air. GM chose to make the air removal process automatic, so why not take advantage of it? GM will not remove the air bleed tubes because they are needed. The routing through the TB on Gen III engines was just convenient. once they found out that heating the TB was not necessary, they removed the tube from the TB. They did not remove the air bleed tubes because they are necessary.
Regards, John McGraw